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Dr Todd R Yokley

Central Classroom 106R

Personal Biography Statement

Dr. Yokley is a biological anthropologist whose research involves analyzing how climate and other selective pressures have shaped human evolution. To date, he has concentrated primarily on variation in the nose and nasal passages, which he has measured through a combination of traditional skeletal measurements and soft-tissue measurements derived from clinical CT scans. This research has revealed a pattern of variation in human nasal passage structure that appears to reflect selection for more efficient heat and moisture exchange under certain environmental conditions. Dr. Yokley is also interested in broader issues such as bioenergetics of past human populations, the emergence of modern humans, climatic adaptation among non-human mammals, and primate paleontology.

Listen to Dr. Yokley talk about his research on Colorado Matters.

Educational Biography

Ph.D. in Biological Anthropology and Anatomy, 2006, Duke University.
M.A. in Anthropology, 1999, Northern Illinois University.
B.S. in Zoology (Anthropology Minor), 1996, University of Tennessee.

Selected Publications

Maddux SD, Butaric LN, Yokley TR, and Franciscus RG. (2017). Ecogeographic variation across morphofunctional units of the human nose. American Journal of Physical Anthropology 162, 103-119. Link to Article

Maddux SD, Yokley TR, Svoma BM, and Franciscus RG. (2016). Absolute humidity and the human nose: A reanalysis of climate zones and their influence on nasal form and function. American Journal of Physical Anthropology 161, 309-320. Link to Article

Holton NE, Alsamawi A, Yokley TR, and Froehle AW. (2016). The ontogeny of nasal shape: An analysis of sexual dimorphism in a longitudinal sample. American Journal of Physical Anthropology 160, 52-61. Link to Article

Hartman CH, Holton NE, Miller SF, Yokley TR, Marshall S, Srinivasan S, and Southard TE. (2016). Nasal Septal Deviation and Facial Skeletal Asymmetries. The Anatomical Record 299, 295-306. Link to Article

Holton NE, Yokley TR, Froehle AW, and Southard TE. (2014). Ontogenetic scaling of the human nose in a longitudinal sample: Implications for genus Homo facial evolution. American Journal of Physical Anthropology 153, 52-60. Link to Article

Selected Publications

Froehle AW, Yokley TR, and Churchill SE. (2013). Energetics and the origin of modern humans. In: Smith FH and Ahern JCM, editors, The Origins of Modern Humans: Biology Reconsidered, Second Edition. Hoboken: Wiley-Blackwell, pp. 285-320. Link to Book

Holton NE, Yokley TR, and Butaric LN. (2013). The morphological interaction between the nasal cavity and maxillary sinuses in living humans. The Anatomical Record 296: 414-426. Link to Article

Holton NE, Yokley TR, and Figueroa A (2012). Nasal septal and craniofacial form in European- and African-derived populations. Journal of Anatomy 221, 263-274. Link to Article

Holton NE, Yokley TR, and Franciscus RG. (2011). Climatic adaptation and Neandertal facial evolution: a comment on Rae et al. (2011). Journal of Human Evolution 61, 624-627. Link to Article

Yokley TR. (2009). Ecogeographic variation in human nasal passages. American Journal of Physical Anthropology 138, 11-22. Link to Article

Courses Taught


Office Hours

Monday - [13:00 to 15:30]
Wednesday - [13:00 to 15:30]

Photo of Dr Todd R Yokley

Current Semester Schedule

43092 ANT-49AE-001 Wyoming Paleontology TBA TBA-TBA

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